amber inclusion archive

Spiders and Scorpions

A small but well preserved and well postured spider in amber.
Spider in amber

A large spider in amber

Spider in amber

Amblypygi Looks like a spider but look closer and you see a whiptail scorpion in amber.

Spider whiptail scorpion in amber

This one has a double-tail pseudo-scorpion in amber

Double whiptail in amber

This one mite be a spider

Spider mite in amber

Spiders were good caring parents

Spider with eggs in amber

CHELICERATA (1c, 13o., 67f., 96g., 146sp.)

Arachnida (13o., 67f., 96g., 146sp.)

Amblypygi (1g., 3sp.)

Family incertae sedis

Kronocharon engeli Wunderlich, 2015

Kronocharon longicalcaris Wunderlich, 2015

Kronocharon prendinii Engel & Grimaldi, 2014

Araneae (31f., 69g., 105sp.)

Archaeidae

Burmesarchaea grimaldii (Penney, 2003)

Eomysmauchenius septentrionalis Wunderlich, 2008

Filiauchenius paucidentatus Wunderlich, 2008

Lacunauchenius speciosus Wunderlich, 2008

Lacunauchenius longissipes Wunderlich, 2015

Lacunauchenius pilosus Wunderlich, 2015

Planarchaea kopp Wunderlich, 2015

†Burmascutidae

Burmascutum aenigma Wunderlich, 2008

Corinnidae (=Myrmeciidae)?

Dipluridae

Phyxioschemoides collembola Wunderlich, 2015

†Eopsilodercidae

Eopsiloderces loxosceloides Wunderlich, 2008

Eopsiloderces serenitas Wunderlich, 2015

†Fossilcalcaridae

Fossilcalcar praeteritus Wunderlich, 2015

Hersilidae

Burmesiola cretacea Wunderlich, 2011

Burmesiola daviesi Wunderlich, 2015

Spinasilia dissoluta Wunderlich, 2015

†Lagonomegopidae

Archaelagonops propinquus Wunderlich, 2015

Archaelagonops salticoides Wunderlich, 2012

Archaelagonops scorsum Wunderlich, 2015

Burlagonomegops eskovi Penney, 2005

Cymbiolaganops cymbiocalcar Wunderlich, 2015

Lagonoburmops plumosus Wunderlich, 2012

?Lagonomegops tuber Wunderlich, 2015

Lineaburmops beigeli Wunderlich, 2015

Lineaburmops hirsutipes Wunderlich, 2015

Myanlagonops gracilipes Wunderlich, 2012

Parviburmops brevipalpus Wunderlich, 2015

?Paxillomegops brevipes Wunderlich, 2015

Paxillomegops longipes Wunderlich, 2015

Picturmegops signatus Wunderlich, 2015

Leptonetidae

Palaeoleptoneta calcar Wunderlich, 2012

Liphistidae

Cretaceothele lata Wunderlich, 2015

†Micropalpimanidae

Micropalpimanus poinari Wunderlich, 2008

†Mongolarachnidae

Longissipalpus magnus Wunderlich, 2015

Longissipalpus maior Wunderlich, 2015

Longissipalpus minor Wunderlich, 2015

Pedipalparaneus seldeni Wunderlich, 2015

Mysmenidae?

Nephilidae?

‘Nephila’ burmanica Poinar & Buckley, 2011

Oecobiidae

Retrooecobius chomskyi Wunderlich, 2015

Retrooecobius convexus Wunderlich, 2015

Zamilia aculeopectens Wunderlich, 2015

Zamilia antecessor Wunderlich, 2008

Zamilia quattuormammillae Wunderlich, 2015

Oonopidae

Burmorchestina pulcher Wunderlich, 2008

†Pholcochyroceridae

?Pholcochyrocer baculum Wunderlich, 2012

Pholcochyrocer guttulaeque Wunderlich, 2008

Pholcochyrocer pecten Wunderlich, 2012

Spinicreber antiquus Wunderlich, 2015

Spinipalpus vetus Wunderlich, 2015

†Plumorsolidae

Burmorsolus crassus Wunderlich, 2015

Burmorsolus nonplumosus Wunderlich, 2015

†Praeterleptonetidae

Autotomiana hirsutipes Wunderlich, 2015

Biapophyses beate Wunderlich, 2015

Crassitibia longispina Wunderlich, 2015

Crassitibia tenuimana Wunderlich, 2015

Curvitibia curima Wunderlich, 2015

Groehnianus burmensis Wunderlich, 2015

Hypotheridiosoma falcata Wunderlich, 2015

Hypotheridiosoma paracymbium Wunderlich, 2012

Palaeohygropoda myanmarensis Penney, 2004

Parvispina tibialis Wunderlich, 2011

Praeterleptoneta spinipes Wunderlich, 2008

Spinipalpitibia maior Wunderlich, 2015

Psilodercidae

Leclercera ellenbergeri Wunderlich, 2015

Leclercera longissipes Wunderlich, 2012

Leclercera sexaculeata Wunderlich, 2015

Leclercera spicula Wunderlich, 2012

Proterpsiloderces longisetae Wunderlich, 2015

?Psiloderces filiformis Wunderlich, 2012

†Salticoididae

Burmadictyna clava Wunderlich, 2015

Burmadictyna excavata Wunderlich, 2015

Burmadictyna pectin Wunderlich, 2008

Segestriidae

Denticulsegestria rugosa Wunderlich, 2015

Myansegestria caederens Wunderlich, 2015

Myansegestria engin Wunderlich, 2015

Parvosegestria longitibialis Wunderlich, 2015

Parvosegestria obscura Wunderlich, 2015

Parvosegestria pintgu Wunderlich, 2015

Parvosegestria triplex Wunderlich, 2015

Sparassidae (=Eusparassidae)?

†Spatiatoridae

Spatiator putescens Wunderlich, 2015

Vetiator gracilipes Wunderlich, 2015

Telemidae?

Tetrablemmidae

Bicornoculus levis Wunderlich, 2015

Electroblemma bifida Selden, Zhang & Ren, 2016

?Eogamasomorpha clara Wunderlich, 2015

Eogamasomorpha nubila Wunderlich, 2008

Eoscaphiella ohlhoffi Wunderlich, 2011

Praeterpaculla armatura Wunderlich, 2015

Praeterpaculla biacuta Wunderlich, 2015

Praeterpaculla dissolata Wunderlich, 2015

Praeterpaculla equester Wunderlich, 2015

Praeterpaculla tuberosa Wunderlich, 2015

Saetosoma filiembolus Wunderlich, 2012

Uniscutosoma aberrans Wunderlich, 2015

Tetragnathidae?

Thomisidae?

Theridiosomatidae

Leviunguis bruckschi Wunderlich, 2012

Theridiidae

Cretotheridion inopinatum Wunderlich, 2015

Uloboridae

Bicalamistrum mixtum Wunderlich, 2015

Burmuloborus antefixus Wunderlich, 2015

Burmuloborus parvus Wunderlich, 2008

?Burmuloborus prolongatus Wunderlich, 2015

Microuloborus birmanicus Wunderlich, 2015

Ocululoborus curvatus Wunderlich, 2012

Palaeomiagrammopes vesica Wunderlich, 2008

Paramiagrammopes cretaceus Wunderlich, 2008

Paramiagrammopes longiclypeus Wunderlich, 2015

Paramiagrammopes patellidens Wunderlich, 2015

Family incertae sedis

Furcembolus andersoni Wunderlich, 2008

Bartel, C., Dunlop, J.A. & Bird, T.L. 2016. The second camel spider (Arachnida, Solifugae)

Burmese amber. Arachnology, 17(3), 161-164.

Cai, C. & Huang, D. 2016c (on-line). A new genus of whip-scorpions in Upper Cretaceous Burmese amber: Earliest fossil record of the extant subfamily Thelyphoninae (Arachnida: Thelyphonida: Thelyphonidae. Cretaceous Research, 69, 100-105.

Dunlop, J.A., Bird, T.L., Brookhart, J.O. & Bechly, G. 2015. A camel spider from Cretaceous Burmese amber. Cretaceous Research, 56, 265-273.

Dunlop, J.A. & Mitov, P.G. 2011. The first fossil cyphophthalmid harvestman from Baltic amber. Arachnologische Mitteilungen, 40, 47-54.

Engel, M.S. & Grimaldi, D.A. 2014b. Whipspiders (Arachnida: Amblypygi) in amber from the Early Eocene and mid-Cretaceous, including maternal care. Novitates Paleoentomologicae, No. 9, 17pp.

Lourenço, W.R. 2015a. A new subfamily, genus and species of fossil Scorpiones from Cretaceous Burmese amber (Scorpiones: Palaeoeuscorpiidae). In Wunderlich, J. (ed.) Mesozoic Spiders (Araneae). Beiträge zur Araneologie, 9, 457-464.

Lourenço, W.R. 2015b. Clarification of the familiar status of the genus Palaeoburmesebuthus Lourenço, 2002 from Cretaceous Burmese amber (Scorpiones: Archaeobuthidae: Palaeoburmesebuthinae). In Wunderlich, J. (ed.) Mesozoic Spiders (Araneae). Beiträge zur Araneologie, 9, 465-475.

Lourenço, W.R. 2015c. New contribution to the knowledge of Cretaceous Burmese amber scorpions: descriptions of two new species of BetaburmesebuthusLourenço, 2015 (Scorpiones: Archaeobuthidae: Palaeoburmesebuthinae). Arachnida, Rivista Aracnologica Italiana, 3, 27–36.

Lourenço, W.R. 2015d. An unusual new species of Chaerilobuthus Lourenço & Beigel, 2011 (Scorpiones: Chaerilobuthidae) from the Cretaceous amber of Myanmar (Burma). Arachnida, Rivista Aracnologica Italiana, 5, 44–48.

Lourenço, W.R. 2016a. A new genus and three new species of scorpions from Cretaceous

Burmese amber (Scorpiones: Chaerilobuthidae: Palaeoeuscorpiidae. Arthropoda Selecta, 25(1), 67-74.

Lourenço, W.R. 2016b. A preliminary synopsis on amber scorpions with special reference to Burmite species: an extraordinary development of our knowledge in only 20 years. ZooKeys, 600, 75-87.

Lourenço, W.R. & Beigel, A. 2011. A new scorpion fossil from the Cretaceous amber of Myanmar (Burma). New phylogenetic implications. Comptes Rendus Palevol, 10, 635-639.

Lourenço, W.R. & Beigel, A. 2015. A new genus and species of Palaeoburmesebuthinae Lourenço, 2014 (Scorpiones: Archaeobuthidae) from Cretaceous amber of Myanmar. In Wunderlich, J. (ed.) Mesozoic Spiders (Araneae). Beiträge zur Araneologie, 9, 465-475.

Lourenço, W.R. & Velten J. 2015. Another new species of Chaerilobuthus Lourenço & Beigel, 2011 (Scorpiones: Chaerilobuthidae) from the Cretaceous amber of Myanmar (Burma). Arachnida, Rivista Aracnologica Italiana, 5, 2–8.

Lourenço W.R. & Velten, J. 2016a. One more new species of Betaburmesebuthus Lourenço, 2015

(Scorpiones: Palaeoburmesebuthinae) from Cretaceous burmite. Arachnida, Rivista Aracnologica Italiana, 6, 4–11.

Lourenço W.R. & Velten, J. 2016b A new genus and species of fossil scorpion from Burmese Cretaceous amber (Scorpiones: Buthoidea: Buthidae. Arachnida, Rivista Aracnologica Italiana, 10, 2-9.

Lourenço W.R. & Velten, J. 2016c A sixth new species of Cretaceous Burmese amber scorpion of the genus Betaburmesebuthus Lourenço, 2015 (Scorpiones: Palaeoburmesebuthidae. Arachnida, Rivista Aracnologica Italiana, 10, 10-17.

Dunlop, J.A. & Penney, D. 2012. Fossil Arachnids. Siri Scientific Press, Manchester. 192pp.

Dunlop, J.A., Selden, P.A. & Giribet, G. 2016. Penis morphology in a Burmese amber harvestman. Science of Nature, 103(11), 1-5.

Penney, D. 2003. Afrarchaea grimaldii, a new species of Archaeidae (Araneae) in Cretaceous Burmese amber. Journal of Arachnology, 31, 122-130.

Penney, D. 2004. A new genus and species of Pisauridae (Araneae) in Cretaceous Burmese amber. Journal of Systematic Palaeontology, 2(2), 141-145.

Penney, D. 2005. The fossil spider family Lagonomegopidae in Cretaceous ambers with descriptions of a new genus and species from Myanmar. Journal of Arachnology, 33, 439-444.

Penny, D. 2006. Fossil oonopid spiders in Cretaceous ambers from Canada and Myanmar. Palaeontology, 49(1), 229-235.

Penney, D. 2012. Predatory behaviour of Cretaceous social orb-weaving spiders: comment. Historical Biology, 26 (1), 132-134. [Published on-line 2012, printed 2014]

Poinar, G.Jr & Buckley, R. 2011 (on-line). Predatory behaviour of the social orb-weaver spider, Geratonephila burmanica n. gen., n. sp. (Araneae: Nephilidae) with its wasp prey, Cascoscelio incassus n. gen. n. sp. (Hymenoptera: Platygastridae) in Early Cretaceous Burmese amber. Historical Biology, 24 (5), 519-525.

Poinar, G.Jr & Buckley, R. 2012 (on-line). Predatory behaviour of the social orb-weaver spiders: response to Penney. Historical Biology, 26 (1), 135-136.

>Wunderlich, J. 2011. (ed.) Extant and fossil spiders (Araneae). Beiträge zur Araneologie, 6,

[not checked]

Wunderlich, J. 2012a. On the fossil spider (Araneae) fauna in Cretaceous ambers, with descriptions of new taxa from Myanmar (Burma) and Jordan, and on the realtionships of the Superfamily Leptonetoidea. In. Wunderlich, J. (ed.) Fifteen papers on extant and fossil spiders (Araneae). Beiträge zur Araneologie, 7, 157-232.

Wunderlich, J. 2012b. Description of the first fossil Ricinulei in amber from Burma (Myanmar), the first report of this arachnid order from the Mesozoic and from Asia, with notes on the related extinct Order Trigonotarbida. In. Wunderlich, J. (ed.) Fifteen papers on extant and fossil spiders (Araneae). Beiträge zur Araneologie, 7, 233-244.

Wunderlich, J. 2015a. “Frozen behaviour”: The oldest fossil proofs of spider eating spiders (Araneae) in 45 million years old Eocene Baltic amber and in 100 million years old Cretaceous amber from Myanmar (Burma). In: Wunderlich, J. (ed.) Mesozoic Spiders (Araneae). Beiträge zur Araneologie, 9, 15-20.

Wunderlich, J. 2015b. On the evolution and the classification of spiders, the Mesozoic spider faunas, and descriptions of new Cretaceous taxa mainly in amber from Myanmar (Burma) (Arachnida:

Araneae). In: Wunderlich, J. (ed.) Mesozoic Spiders (Araneae). Beiträge zur Araneologie, 9, 21-408.

Wunderlich, J. 2015c. New and rare Arachnida in Cretaceous Burmese amber (Amblypygi, Ricinulei and Uropygi: Thelephonida). In: Wunderlich, J. (ed.) Mesozoic Spiders (Araneae). Beiträge zur Araneologie, 9, 409-436.

Selden, P.A. & Zhang, W. & Ren, D. 2016. A bizarre armoured spider (Araneae: Tetrablemmidae) from Upper Cretaceous Myanmar amber. Cretaceous Research, 66, 129-135.

Wunderlich, J. 2008a. Descriptions of fossil spider (Araneae) taxa mainly in Baltic amber, as well as on certain related extant taxa. In: Fossil and extant spiders (Araneae). Beitraege zur Araneologie, 5, 44-139.

Wunderlich, J. 2008b. The dominance of ancient spider families of the Araneae: Haplogynae in the Cretaceous, and the late diversification of advanced ecribellate spiders of the Entelegynae after the Cretaceous-Tertiary boundary extinction events, with descriptions of new families. In: Fossil and extant spiders (Araneae). Beitraege zur Araneologie, 5, 524-675.

Selden, P.A., Dunlop, J.A., Giribet, G., Zhang, W. & Ren, D. 2016. The oldest armoured harvestman (Arachnida: Opiliones: Laniatores), from Upper Cretaceous Myanmar amber. Cretaceous Research, 65, 206-212.

Santiago-Blay, J. A., V. Fet, M. E. Soleglad, S. & Anderson. 2004. A new genus and subfamily of scorpions from Cretaceous Burmese amber (Scorpiones: Chaerilidae). Revista Ibérica de Aracnología, 9, 3-14.

Santiago-Blay, J.A., Fet, V., Soleglad, M.E. & Craig, P.R. 2004. The second Cretaceous scorpion specimen from Burmese amber (Arachnida: Scorpiones). Journal of Systematic Palaeontology, 2(2), 147-152.

Rossi, A. 2015. A new family, genus and species of scorpion from the burmite of Myanmar (Scorpiones: Sucinlourencoidae). Rivista Aracnologica Italiana, 1, 3-21.

Poinar, G.O.Jr. 2008a. Palaeosiro burmanicum n. gen., n. sp., a fossil Cyphophthalmi (Arachnida: Opiliones: Sironidae) in Early Cretaceous Burmese amber. In: Makarov, S.E. & Dimitrijević, R.N. (eds.) Advances in Arachnology and Developmental Biology. Institute of Zoology, University of Belgrade, Monographs, 12, 267-274.

Judson, M.L.I. 2000. Electrobisium acutum Cockerell, a cheiridiid pseudoscorpion from Burmese amber, with remarks on the validity of the Cheiridioidea (Arachnida, Chelonethi). Bulletin of the Natural History Museum, Geology Series, 56(1), 79-83.

Lourenço, W.R. 2002. The first scorpion fossil from the Cretaceous amber of Myanmar (Burma). New implications for the phylogeny of Buthoidea. Comptes Rendus Palevol, 1, 97-101.

Lourenço, W.R. 2012. About the scorpion fossils from the Cretaceous amber of Myanmar (Burma) with the descriptions of a new family, genus and species. Acta Biológica Paranaense, Curitiba, 41 (3-4), 75-87.

Lourenço, W.R. 2013. A new species of Chaerilobuthus Lourenço & Beigel, 2011 from Cretaceous Burmese amber (Scorpiones: Chaerilobuthidae). Acta Biológica Paranaense, Curitiba, 42(1-2), 1–5.

Engel, M.S., Breitkreuz, L.C.V., Cai, C., Alvarado, M., Azar, D. & Huang, D. 2016. The first Mesozoic microwhip scorpion (Palpigradi): a new genus and species in mid-Cretaceous amber from Myanmar. Science of Nature, 103(19), 1-7.

Judson, M.L.I. 2009. Cheliferoid pseudoscorpions (Arachnida, Chelonethi) from the Lower Cretaceous of France. Geodiversitas, 31(1), 61-71.

Our taxa list is based on the amazing work of Dr Andrew J. Ross who has prepared a newer updated list and kindly offers it for free download
Amber lovers and Paleos can dig up the latest updated version here:
http://www.nms.ac.uk/media/1154465/burmese-amber-taxa-v2017_2.pdf